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Heller will no longer find 'Events' throughout NYZ, instead starting side-missions by hacking into Blacknet. In prototype 2 Heller can control a pack of Brawlers (similar to Prototype '​s Hunters) to attack anything he desires (note: this power is limited to main variant of Brawler).

Consuming has not changed since the original Prototype, with the exception of some enemies, which are consumed upon grabbing them (e.g. There are several other ways to kill enemies, ranging from throwing a car at a group of Blackwatch soldiers to hacking a mutant's head or using super powers.

Players will be able to dismember enemies, a force that will become more useful as the game advances. Tendrils sprout from Heller's arm and has a variety of purposes.Tendrils can be used to smash objects into other objects, such as a car into a tank, making use of the special 'Black Hole' attack. To give gamers more power in the game, the developers have added tendrils( blade like structure in the hands of Heller). Heller will also undergo mutations and upgrades that let players decide how they want to play as Heller. The sonar detector includes a new pulse ability that has the feature to make it easier for the player to find people hiding, instead of looking in a large crowd for a person with an icon above their head indicating the target. Heller also has strength and agility of that of a superhuman, near-invulnerability to harm, near-flight leaping and gliding, he also has infinite stamina, has great speed, and also has a sonar sense. James Heller can also sneak up on unsuspecting human enemies, inject the Blacklight virus in them turning them into "BiologicalBomb" to blast(blow up). Heller will be able to use weapons in the game, such as ripping the Gatling cannon off a tank and using it against enemies. To make sure player do not become overwhelmed by the enemies, Radical has created a dodging system and new, more realistic AI. Due to Blackwatch's actions in the Yellow Zone, if the player assumes the role of a soldier, people will react to him in a way that shows that they want nothing to do with him.

Taking people's identities and changing shape into them has become more tactical. James Heller the protagonist of the game has many super ultra powers which include: he can shift his body and assume the identity of other people and also their memories by feeding on their mind. The game was a best seller for a few years. The is also of a revenge type, as Heller wants to kill Alex Mercer, hero of the original Prototype, as his family was killed due to the outbreak of the Backlight virus created by Alex Mercer. The game features a new protagonist, James Heller, as he goes on a quest to destroy the deadly Backlight virus.